OUR CHARMING TETRAPODS

The irresistable tetrapod Tiktaalik

In the late 1930s, our understanding of the transition of fish to tetrapods — and the eventual jump to modern vertebrates — took an unexpected leap forward. 

The evolutionary a'ha came from a single partial fossil skull found on the shores of a riverbank in Eastern Canada. 

Meet the Stegocephalian, Elpistostege watsoni, an extinct genus of finned tetrapodomorphs that lived during the Late Givetian to Early Frasnian of the Late Devonian — 382 million years ago. 

Elpistostege watsoni — perhaps the sister taxon of all other tetrapods — was first described in 1938 by British palaeontologist and elected Fellow of the Royal Society of London, Thomas Stanley Westoll. Westroll was an interesting fellow whose research interests were wide-ranging. He was a vertebrate palaeontologist and geologist best known for his innovative work on Palaeozoic fishes and their relationships with tetrapods. 

Elpistostege watsoni

As a specialist in early fish, Westoll was the perfect person to ask to interpret that single partial skull roof discovered at the Escuminac Formation in Quebec, Canada. 

His findings and subsequent publication named Elpistostege watsoni and helped us to better understand the evolution of fishes to tetrapods — four-limbed vertebrates — one of the most important transformations in vertebrate evolution. 

Hypotheses of tetrapod origins rely heavily on the anatomy of but a few tetrapod-like fish fossils from the Middle and Late Devonian, 393–359 million years ago. 

These taxa — known as elpistostegalians — include Panderichthys, Elpistostege and Tiktaalik — none of which had yet to reveal the complete skeletal anatomy of the pectoral fin. 

Elpistostege watsoni

None until 2010 that is, when a complete 1.57-metre-long articulated specimen was found and described by Richard Cloutier et al. in 2020. 

The specimen helped us to understand the origin of the vertebrate hand. Stripped from its encasing stone, it revealed a set of paired fins of Elpistostege containing bones homologous to the phalanges (finger bones) of modern tetrapods and is the most basal tetrapodomorph known to possess them. 

Once the phalanges were uncovered, prep work began on the fins. The fins were covered in wee scales and lepidotrichia (fin rays). The work was tiresome, taking more than 2,700 hours of preparation but the results were thrilling. 

Origin of the Vertebrate Hand

We could now clearly see that the skeleton of the pectoral fin has four proximodistal rows of radials — two of which include branched carpals — as well as two distal rows organized as digits and putative digits. 

Despite this skeletal pattern — which represents the most tetrapod-like arrangement of bones found in a pectoral fin to date blurring the line between fish and land vertebrates — the fin retained lepidotrichia (those wee fin rays) distal to the radials. 

This arrangement confirmed an age-old question — showing us for the first time that the origin of phalanges preceded the loss of fin rays, not the other way around.

E. watsoni is very closely related to Tiktaalik roseae found in 2004 in the Canadian Arctic — a tetrapodomorpha species also known as a Choanata. These were advanced forms transitional between fish and the early labyrinthodonts playfully referred to as fishapods — half-fish, half-tetrapod in appearance and limb morphology. 

Up to that point, the relationship of limbed vertebrates (tetrapods) to lobe-finned fish (sarcopterygians) was well known, but the origin of major tetrapod features remained obscure for lack of fossils that document the sequence of evolutionary changes — until Tiktaalik. While Tiktaalik is technically a fish, this fellow is as far from fish-like you can be and still be a card-carrying member of the group. 

Tiktaalik roseae

Complete with scales and gills, this proto-fish lacked the conical head we see in modern fish but had a rather flattened triangular head more like that of a crocodile. 

Tiktaalik had scales on its back and fins with fin webbing but like early land-living animals, it had a distinctive flat head and neck. He was a brawny brute. The shape of his skull and shoulder look part fish and part amphibian.

The watershed moment came as Tiktaalik was prepped. Inside Tiktaalik's fins, we find bones that correspond to the upper arm, forearm and even parts of the wrist — all inside a fin with webbing — remarkable! 

Its fins have thin ray bones for paddling like most fish, but with brawny interior bones that gave Tiktaalik the ability to prop itself up, using his limbs for support. I picture him propped up on one paddle saying, "how you doing?" 

Six years after Tiktaalik was discovered by Neil Shubin and team in the ice-covered tundra of the Canadian Arctic on southern Ellesmere Island, a team working the outcrops at Miguasha on the Gaspé Peninsula discovered the only fully specimen of E. watsoni found to date — greatly increasing our knowledge of this finned tantalizingly transitional tetrapodomorph. 

E. watsoni fossils are rare — this was the fourth specimen collected in over 130 years of hunting. Charmingly, the specimen was right on our doorstop — extracted but a few feet away from the main stairs descending onto the beach of Miguasha National Park. 

L'nu Mi’gmaq First Nations of the Gespe’gewa’gi Region

Miguasha is nestled in the Gaspésie or Gespe’gewa’gi region of Canada — home to the Mi’gmaq First Nations who self-refer as L’nu or Lnu. The word Mi’gmaq or Mi’kmaq means the family or my allies/friends in Mi'kmaw, their native tongue (and soon to be Nova Scotia's provincial first language). They are the people of the sea and the original inhabitants of Atlantic Canada having lived here for more than 10,000 years. 

The L'nu were the first First Nation people to establish contact and trade with European explorers in the 16th and 17th centuries — and perhaps the Norse as early as the turn of the Millenium. Sailing vessels filled with French, British, Scottish, Irish and others arrived one by one to lay claim to the region — settling and fighting over the land. As each group rolled out their machinations of discovery, tensions turned to an all-out war with the British and French going head to head. I'll spare you the sordid details but for everyone caught in the crossfire, it went poorly.

North America Map 1775 (Click to Enlarge)

Cut to 1760, the British tipped the balance with their win at the Battle of the Restigouche, the last naval battle between France and England for possession of the North American continent — Turtle Island. 

The bittersweet British victory sparked the American War of Independence. 

For the next twenty years, the L'nu would witness and become embroiled in yet another war for these lands, their lands — first as bystanders, then as American allies, then intimidated into submission by the British Royal Navy with a show of force by way of a thirty-four gun man-of-war, encouraging L'nu compliance — finally culminating in an end to the hostilities with the 1783 Treaty of Paris. 

The peace accord held no provisions for the L'nu, Métis and First Nations impacted. None of these newcomers was Mi'kmaq — neither friends nor allies.

It was to this area some sixty years later that the newly formed Geological Survey of Canada (GSC) began exploring and mapping the newly formed United Province of Canada. Geologists in the New Brunswick Geology Branch traipsed through the rugged countryside that would become a Canadian province in 1867. 

It was on one of these expeditions that the Miguasha fossil outcrops were discovered. They, too, would transform in time to become Miguasha National Park or Parc de Miguasha, but at first, they were simply the promising sedimentary exposures on the hillside across the water —  a treasure trove of  Late Devonian fauna waiting to be discovered.

In the summer of 1842, Abraham Gesner, New Brunswick’s first Provincial Geologist, crossed the northern part of the region exploring for coal. Well, mostly looking for coal. Gesner also had a keen eye for fossils and his trip to the Gaspé Peninsula came fast on the heels of a jaunt along the rocky beaches of Chignecto Bay at the head of the Bay of Fundy and home to the standing fossil trees of the Joggins Fossil Cliffs. 

Passionate about geology and chemistry, he is perhaps most famous for his invention of the process to distil the combustible hydrocarbon kerosene from coal oil — a subject on which his long walks exploring a budding Canada gave him a great deal of time to consider. We have Gesner to thank for the modern petroleum industry. He filed many patents for clever ways to distil the soft tar-like coal or bitumen still in use today.

He was skilled in a broad range of scientific disciplines — being a geologist, palaeontologist, physician, chemist, anatomist and naturalist — a brass tacks geek to his core. Gesner explored the coal exposures and fossil outcrops across the famed area that witnessed the region become part of England and not France — and no longer L'nu.

Following the Restigouche River in New Brunswick through the Dalhousie region, Gesner navigated through the estuary to reach the southern coast of the Gaspé Peninsula into what would become the southeastern coast of Quebec to get a better look at the cliffs across the water. He was the first geologist to lay eyes on the Escuminac Formation and its fossils.

In his 1843 report to the Geologic Survey, he wrote, “...I found the shore lined with a coarse conglomerate. Farther eastward the rocks are light blue sandstones and shales, containing the remains of vegetables. (...) In these sandstone and shales, I found the remains of fish and a small species of tortoise with fossil foot-marks.”

We now know that this little tortoise was the famous Bothriolepis, an antiarch placoderm fish. It was also the first formal mention of the Miguasha fauna in our scientific literature. Despite the circulation of his report, Gesner’s discovery was all but ignored — the cliffs and their fossil bounty abandoned for decades to come. Geologists like Ells, Foord and Weston, and the research of Whiteaves and Dawson, would eventually follow in Gesner's footsteps.

North America Map 1866 (Click to Enlarge)

Over the past 180 years, this Devonian site has yielded a wonderfully diverse aquatic assemblage from the Age of Fishes — five of the six fossil fish groups associated with the Devonian including exceptionally well-preserved fossil specimens of the lobe-finned fishes. 

This is exciting as it is the lobe-finned fishes — the sarcopterygians — that gave rise to the first four-legged, air-breathing terrestrial vertebrates – the tetrapods. 

Fossil specimens from Miguasha include twenty species of lower vertebrates — anaspids, osteostra-cans, placoderms, acanthodians, actinopterygians and sarcopterygians — plus a limited invertebrate assemblage, along with terrestrial plants, scorpions and millipedes.

Originally interpreted as a freshwater lacustrine environment, recent paleontological, taphonomic, sedimentological and geochemical evidence corroborates a brackish estuarine setting — and definitely not the deep waters of the sea. This is important because the species that gave rise to our land-living animals began life in shallow streams and lakes. It tells us a bit about how our dear Elpistostege watsoni liked to live — preferring to lollygag in cool river waters where seawater mixed with fresh. Not fully freshwater, but a wee bit of salinity to add flavour.  

• Photos: Elpistostege watsoni (Westoll, 1938 ), Upper Devonian (Frasnian), Escuminac formation, Parc de Miguasha, Baie des Chaleurs, Gaspé, Québec, Canada. John Fam, VanPS
• Origin of the Vertebrate Hand Illustration, https://www.nature.com/articles/s41586-020-2100-8
• Tiktaalik Illustration: By Obsidian Soul - Own work, CC BY 4.0, https://commons.wikimedia.org/w/index.php?curid=47401797

References & further reading:

• From Water to Land: https://www.miguasha.ca/mig-en/the_first_discoveries.php
• UNESCO Miguasha National Park: https://whc.unesco.org/en/list/686/
• Office of L'nu Affairs: https://novascotia.ca/abor/aboriginal-people/
• Cloutier, R., Clement, A.M., Lee, M.S.Y. et al. Elpistostege and the origin of the vertebrate hand. Nature 579, 549–554 (2020). https://doi.org/10.1038/s41586-020-2100-8
• Daeschler, E. B., Shubin, N. H. & Jenkins, F. A. Jr. A Devonian tetrapod-like fish and the evolution of the tetrapod body plan. Nature 440, 757–763 (2006).
• Shubin, Neil. Your Inner Fish: A Journey into the 3.5 Billion History of the Human Body.
• Evidence for European presence in the Americas in AD 1021: https://www.nature.com/articles/s41586-021-03972-8

VANCOUVER ISLAND'S HASLAM FORMATION

Steller's Jay, Cyanocitta stelleri

One of the classic Vancouver Island fossil localities is the Santonian-Maastrichtian, Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake, Nanaimo, British Columbia, Canada.

The quarry is no longer active as such though there is a busy little gravel quarry a little way down the road closer to Ammonite falls near Benson Creek Falls.

Today it is an active motocross site and remains one of the classic localities of the Nanaimo Group. We find well-preserved nautiloids and ammonites — Canadoceras, Pseudoschloenbachia, Epigoniceras — the bivalves — Inoceramus, Sphenoceramus— gastropods, and classic Nanaimo Group decapods — Hoploparia, Linuparus. We also find fossil fruit and seeds which tell the story of the terrestrial history of Vancouver Island.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

It was John Fam, Vice-Chair, Vancouver Island Paleontological Society (VanPS), who originally told me about the locality. John is one of the most delightful and knowledgeable people you'd be well-blessed to meet.

While he lived on Vancouver Island, he was an active member of the VanPS back when I was Chair. Several of the best joint VIPS/VanPS paleontological expeditions were planned with or instigated by his passion for fossils. I tip my hat to him for his passion and shared love of all things paleo.

John grew up 15 minutes from the motocross locality and used to collect there a few times a week with his father. John has wonderful parents and since marrying his childhood sweetheart, the amazing Grace, those excellent genetics, curiosity and love of fossils are now being passed to a new generation. It's lovely to see John and Grace continuing tradition with two boys of their own.

I met John way back then and did an overnight at his parent's house the Friday before a weekend field trip to Jurassic Point. It was a joy to have him walk me through his collections and tell his stories from earlier years. After learning about the site from John, I headed up to the Motocross Pit with my Uncle Doug. He was a delightful man who grew up on the coast and had explored much of it but not the fossil site just 10-minutes from his home. It was wonderful to walk through time with him so many years ago and then again solo this past year with sadness in my belly that one of the best I've ever known has left this Earth.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

There were some no trespassing signs up but no people around, so I walked the periphery looking for the bedrock of the Haslam.

The rocks we find here were laid down south of the equator as small, tropical islands. They rode across the Pacific heading north and slightly east over the past 80 million years to where we find them today.

Jim Haggart and Peter Ward have done much to increase our understanding of the molluscan fauna of the Nanaimo Group. Personally, both personify the charming Indiana Jones school of rugged manly palaeontologists you picture in popular film. Professionally, their singular contributions and collaborative efforts have helped shape our understanding of the correlation of Nanaimo Group fauna to those we find in the Gulf Islands of British Columbia and down in the San Juan Islands of Washington State.

Their work builds on the work of Usher (1952), Matsumoto (1959a, 1959b) and Mallory (1977). A healthy nod goes out to the work of Muller and Jeletzky (1970) for untangling the lithostratigraphic and biostratigraphic foundation for our knowledge of the Nanaimo Group.

Candoceras yokoyama, Photo: John Fam, VanPS

As I walked along the bedrock of the Haslam, a Steller's Jay, Cyanocitta stelleri, followed me from tree to tree making his guttural shook, shook, shook call. Instructive, he seemed to be encouraging me, timing his hoots to the beat of my hammer. Vancouver Island truly has glorious flora and fauna.

Fancy some additional reading? Check out a paper published in the Journal of Paleontology back in 1989 by Haggard and Ward on new Nanaimo Group Ammonites from British Columbia and Washington State.

In it, they look at the ammonite species Puzosia (Mesopuzosia) densicostata Matsumoto, Kitchinites (Neopuzosia) japonicus Spath, Anapachydiscus cf. A. nelchinensis Jones, Menuites cf. M. menu (Forbes), Submortoniceras chicoense (Trask), and Baculites cf. B. boulei Collignon are described from Santonian--Campanian strata of western Canada and northwestern United States.

Stratigraphic occurrences and ranges of the species are summarized and those taxa important for correlation with other areas in the north Pacific region and Late Cretaceous ammonite fauna of the Indo-Pacific region. Here's the link: https://www.jstor.org/stable/1305358?seq=1

Peter Ward is a prolific author, both of scientific papers and more popularized works. I highly recommend his book Gorgon: Paleontology, Obsession, and the Greatest Catastrophe in Earth's History. It is an engaging romp through a decade's research in South Africa's Karoo Desert.

Photo: Candoceras yokoyamai from Upper Cretaceous Haslam formation (Lower Campanian) near Nanaimo, British Columbia. One of the earliest fossils collected by John Fam (1993). Prepared using only a cold chisel and hammer. Photo & collection of John Fam, VIPS.

BLUE JAY: KWASK'WAS

If you live in North American, there is a high probability that you have seen or heard the bird song of the Blue jay, Cyanocitta cristata (Linnaeus, 1758).

Blue Jays are in the family Corvidae — along with crows, ravens, rooks, magpies and jackdaws. They belong to a lineage of birds first seen in the Miocene — 25 million years ago. 

These beautifully plumed, blue, black and white birds can be found across southern Canada down to Florida. The distinctive blue you see in their feathers is a trick of the light. Their pigment, melanin, is actually a rather dull brown. The blue you see is caused by scattering light through modified cells on the surface of the feather as wee barbs.

Blue jays like to dine on nuts, seeds, suet, arthropods and some small vertebrates. 

If you are attempting to lure them to your yard with a bird feeder, they prefer those mounted on trays or posts versus hanging feeders. They will eat most anything you have on offer but sunflower seeds and peanuts are their favourites. 

They have a fondness for acorns and have been credited with helping expand the range of oak trees as the ice melted after the last glacial period.  

Their Binomial name, Cyanocitta cristata means, crested, blue chattering bird. I might have amended that to something less flattering, working in a Latin word or two for shrieks and screams — voce et gemitu or ululo et quiritor. While their plumage is a visual feast, their bird chatter leaves something to be desired. 

In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest and my family, a Blue Jay is known as kwa̱skwa̱s. 

The Kwak’wala word for blue is dzasa and cry is ḵ̕was'id. For interest, the word for bird song in Kwak'wala is t̕sa̱sḵwana. Both their songs and cries are quite helpful if you are an animal living nearby and concerned about predators. 

LYTOCERAS OF THE LIAS

A gorgeous Lytoceras sp. from within the Middle Lias, Lower Jurassic Bridport Sands, Margaritatus Zone, near the secluded town of Eype, Dorset, England. 

Eype is a small village that hugs the English Channel in southwest Dorset near the town of Bridport on the infamous Jurassic Coast. Picture creamy rust-coloured sand, a shelved shingle bank backed by a massive cliff-face and the constant rush of waves.

Aside from the surf, this is a quiet hamlet and a wonderful place to explore along the pebble beach. You'll want to stay well away from the cliffs as rock slides happen quickly and the debris field is much wider than you would think.   

Prepping these lovelies is a tricky business as the flanges are very delicate. You can see that many of the ornamental rings are intact. The specimen itself is amazing but the level of patience and skill that went into the preparation of this lovely cephalopod is world-class. 

Photo: Specimen: 16 cm (just over 6 inches) at its widest point. Craig Chivers, Natural Selection Fossils. If you would like to see more of Craig's work, head on over to his website: naturalselection fossils.com or check out his Instagram account @naturalselectionfossils. You will drool over the incredible ammonites, marine reptile material and some bivalves — plus some paleo art thrown in for good measure. 

NUNATSIRMIUT MOTHER AND CHILD

Nunatsiarmiut Mother and Child, Baffin Island, Nunavut

Warm light bathes this lovely Nunatsiarmiut mother and child from Baffin Island, Nunavut. 

They speak Inuktitut, the mother tongue of the majority of the Nunatsiarmiut who call Baffin Island home. 

Baffin is the largest island in the Arctic Archipelago in the territory of Nunavut in Canada's far north. 

As part of the Qikiqtaaluk Region of Nunavut, Baffin Island is home to a constellation of remote Inuit communities each with a deep cultural connection to the land — Iqaluit, Pond Inlet, Pangnirtung, Clyde River, Arctic Bay, Kimmirut and Nanisivik. 

The ratio of Inuit to non-Inuit here is roughly three to one and perhaps the reason why the Inuktitut language has remained intact and serves as the mother tongue for more than 36,000 residents. Inuktitut has several subdialects — these, along with a myriad of other languages — are spoken across the north.  

If you look at the helpful visual below you begin to get a feel for the diversity of these many tongues. The languages vary by region. There is the Iñupiaq of the Inupiatun/Inupiat; Inuvialuktun of the Inuinnaqtun, Natsilingmiutut, Kivallirmiutut, Aivilingmiutut, Qikiqtaaluk Uannanganii and Siglitun. Kalaallisut is spoken by many Greenlandic peoples though, in northwest Greenland, Inuktun is the language of the Inughuit.

We use the word Inuktitut when referring to a specific dialect and inuktut when referring to all the dialects of Inuktitut and Inuinnaqtun.

Northern Language Map (Click to Enlarge)

Should you travel to the serene glacier-capped wilds and rolling tundra of our far north, you will want to dress for the weather and learn a few of the basics to put your best mukluk shod feet forward. 

The word for hello or welcome in Inuktitut is Atelihai — pronounced ahh-tee-lee-hi. And thank you is nakurmiik, pronounced na-kur-MIIK.  

Perhaps my favourite Inuktitut expression is Naglingniq qaikautigijunnaqtuq maannakautigi, pronounced NAG-ling-niq QAI-kau-ti-gi-jun-naqtuq MAAN-na-KAU-ti-gi. This tongue-twister is well worth the linguistic challenge as it translates to love can travel anywhere in an instant. Indeed it can.

You have likely seen or heard the word Eskimo used in older books to refer to the Inuit, Iñupiat, Kalaallit or Yupik. This misnomer is a colonial term derived from the Montagnais or Innu word ayas̆kimew — netter of snowshoes. It is a bit like meeting a whole new group of people who happen to wear shoes and referring to them all as cobblers — not as a nickname, but as a legal term to describe populations from diverse communities disregarding the way each self-refer. 

Inukshuk / Inuksuk Marker Cairn

For those who identify as Inupiaq or Yupik, the preferred term is Inuit meaning people — though some lingering use of the term Eskimo lives on. The Inuit as a group are made up of many smaller groups. 

The Inuit of Greenland self-refer as Kalaallit or Greenlanders when speaking Kalaallisut. The Inupiat of Alaska, or real people, use Yupik as the singular for real person and yuk to simply mean person.

When taken all together, Inuit is used to mean all the peoples in reference to the Inuit, Iñupiat, Kalaallit and Yupik. Inuit is the plural of inuk or person. 

You likely recognize this word from inuksuk or inukshuk, pronounced ih-nook-suuk — the human-shaped stone cairns built by the Inuit, Iñupiat, Kalaallit, Yupik, and other peoples of the Arctic regions of northern Canada, Greenland, and Alaska — helpful reference markers for hunters and navigation. The word inuksuk means that which acts in the capacity of a human, combining inuk or person and suk, to substitute. 

A World of Confusion

You may be disappointed to learn that our northern friends do not live in igloos. I remember answering the phone as a child and the fellow calling was hoping to speak to my parents about some wonderful new invention perfect for use in an igloo. 

He was disappointed to hear that I was standing in a wooden house with the standard four walls to a room and a handy roof topping it off. "Well, what about your neighbours? Surely some of them live in igloos..." 

It seems that some of the atlases in circulation at the time, and certainly the one he was looking at, simply blanketed everything north of the 49th parallel in a snowy white. His clearly showed an igloo sitting proudly in the centre of the province.

My cousin Shawn brought one such simplified book back from his elementary school in California. British Columbia had a nice image of a grizzly bear and a wee bit further up, a polar bear grinned smugly. British Columbia's beaver population would be sad to know that they did not inhabit the province though there were two chipper beavers with big bright smiles — one in Ontario and another gracing the province of Quebec. Further north, where folk do build igloos, they were curiously lacking. 

Igloos are used for winter hunting trips much the same way we use tents for camping. The Inuit do not have fifty words for snow — you can thank the ethnographer Franz Boas for that wee fabrication — but within the collective languages of the frozen north there are more than fifty words to describe it. And kisses are not nose-to-nose. To give a tender kiss or kunik to a loved one, you press your nose and upper lip to their forehead or cheek and rub gently. 

Fancy trying a wee bit of Inuktitut yourself? This link will bring you to a great place to start: https://inhabitmedia.com/inuitnipingit/

Inuit Language Map:  By Noahedits - Own work, CC BY-SA 4.0. If you want to the image full size, head to this link: https://commons.wikimedia.org/w/index.php?curid=85587388

MANSERICHE BEAUTY: PROLYELLICERAS

This inky beauty is Prolyelliceras ulrichi (Knechtel, 1947) a fast-moving nektonic carnivorous ammonite from Cretaceous lithified, black, carbonaceous limestone outcrops in the Pongo de Manseriche gorge in northwest Peru. 

If you look closely, you can see that this specimen shows a pathology, a slight deviation to the side of the siphonal of the ammonite. We see Prolyelliceras from the Albian to Middle Albian from five localities in Peru.

The canyons of the Amazon River system in the eastern ranges of the Andes of Peru are known by the Indian name pongo. 

The most famous of these is the Pongo de Manseriche, cut by the Marañon River through the eastern range of the Andes, where it emerges from the cordillera into the flat terrane of the Amazon Basin. The fossil exposures here are best explored by boat. The reality of the collecting is similar to the imagined. I was chatting with Betty Franklin, VIPS, about this. They float along and pick up amazing specimen after amazing specimen. When the water rises, the ammonites are aided in their erosion out of the cliffs.  

The Pongo de Manseriche lies nearly 500 miles upstream from Iquitos, and consequently nearly 3,000 miles above the mouth of the Amazon River. It is situated in the heart of the montaña, in a vast region the ownership of which has long been in dispute between Peru and Ecuador, but over which neither country exercises any police or other governmental control. There is an ancient tradition of the indigenous people of the vicinity that one of their gods descended the Marañón and another ascended the Amazon to communicate with him. Together they opened the pass called the Pongo de Manseriche.

Reference: M. M. Knechtel. 1947. Cephalopoda. In: Mesozoic fossils of the Peruvian Andes, Johns Hopkins University Studies in Geology 15:81-139

W. J. Kennedy and H. C. Klinger. 2008. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Lyelliceratinae Spath, 1921. African Natural History 4:57-111. The beauty you see here is in the collection of the deeply awesome José Juárez Ruiz.

CRANBROOK EAGER FORMATION: TUZOIA

This specimen of the arthropod Tuzoia sp. is from the Lower Cambrian Eager Formation of British Columbia. 

Tuzoia is an extinct genus of large bivalved arthropod known from Early to Middle Cambrian marine environments in what is now the Burgess Shale, Emu Bay Shale, Kaili, the Rockslide Formation, the Spence Shale, Wheeler Formation, and Marjum Formation, and the Kinzers Formation. 

As you can see here, they are also found in the Lower Cambrian Eager Formation near Cranbrook British Columbia. 

This particular specimen is from the Rifle Range outcrop near Cranbrook where you can also find numerous fragments and complete specimens of the olenellid trilobites Ollenellus sp. and the larger, more robust Wanneria dunnae, along with Mesonacis eagerensis (Best, 1952).

The site outcrops at a few locations as you head east out of Cranbrook towards Fort Steele. The first trilobites were discovered with the building of the Kootenay Highway connecting Cranbrook to Fort Steele and beyond. Several other localities, including the outcrops at the Silhouette Rife Range — which is literally on a Rifle Range where folks go to shoot at things — is a shade older than the Middle Cambrian Burgess Shale but the fauna here is much less varied. 

The site has been known and collected since the 1920s. Back in the day, fossil collecting was a family affair with folks heading out in their lightly coloured finery to picnic and surface collect the eroding exposures. Cranbrook local, Clement Hungerford Pollen was an engineer and avocational palaeontologist. He promoted collecting the exposures of the Eager Formation around 1921. As a pedigreed Englishman of considerable means, he had invested in the Kootenay Central Railway, revitalizing the town by opening up railway access within the region.